cerda -ced- gismu

x1 is an heir to/is to inherit x2 (object/quality) from x3 according to rule x4.

Pedantically, inheriting an object should be a sumti-raising (tu'a if non-abstract in x2) of inheriting loka ponse the object - the ownership of the object (= posycerda, posyselcerda for unambiguous semantics). See also jgina.

In notes:

x1 is a gene of creature [or locus on creature] x2 determining trait/process x3.
c1 inherits property (right) p1=c2 from c3 according to rule c4.
c2=p1 is the inheritance (legal property) for heir p2=c3, property according to law/custom p3, inherited from c1 according to law/custom c4.
x1 is genetics based on methodology x2.