x1 (partially) occupies/fulfills niche x2 (this is a specific assemblage of roles, purposes, strategies, characteristics, behaviors, physical forms or traits, interactions with an environment, etc.) via particular method/mode/trait x3, in evolutionary or interaction or social or ecological or conceptual or strategic or physical environment x4; the x2 niche is occupied by x1 in way or via form/fashion x3; x2 is a niche in evolutionary system x4.
Not necessarily biological. Care must be taken when discussing niches broadly because abstraction from a specific niche is required. For a subniche or superniche, use "kleniclu". Individual parts or tools of a larger entity (such as wheels on a car or beaks on a bird) could be understood to be valid x1 candidates, but x3 would have to be subparts of them, characteristics of specifically them, or otherwise describe or be related to specifically them (the wheel) rather than the whole (the car); likewise, in multicellular or complex organisms or communal organisms or colonies of organisms, or communities, or societies, or businesses, individual cells or zooids or members of the organization may be said to occupy a niche (be x1) within that structure or organization.